Geography 316:  Biogeography     In progress 12/09/2002

The Biogeography of Bald Eagles (Haliaeetus leucocephalus)
 

by Michael Rochelle, student in Geography 316

Thank you for visiting our site. This web pages was written by a student in Geography 316: Biogeography and edited by the instructor, Barbara Holzman, PhD.  All photos and maps are posted with specific copyright permission for the express use of education on these web pages. The students have tried to be as accurate as possible with the information provided and sources and references are cited at the end of each page.

Species Name: Haliaeetus leucocephalus                 Photo by David and Valerie Peters
                                                                                                 at www.tgrsolution.net/zoo  

Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Falconiformes
Family: Acciptiridae
Genus: Haliaeetus
Species: Haliaeetus leucocephalus


   
  Haliaeetus leucocephalus’ most common name is the bald eagle, although, far from being bald, the taxonomic name literally means white-headed sea-eagle, so bald eagle is a misnomer based upon its appearance (Weidensaul, 1996). The bald eagle represents power, freedom, and for some, spirituality and pristine wildness. It was due to its strong image that Congress officially adopted the bald eagle as the symbol for the United States in 1782, and you will find bald eagle emblems on currency, coins, and the US seal (Stalmaster, 1987). Ironic, that what is revered symbolically was almost brought to the point of extinction in the last century, that our ideologies regarding nature don’t necessarily correspond to our actions that effect it.     

 
Description of Species:
The bald eagle measures from 30 to 43 inches in length, and 70 to 96 inches in width with a wing span from 23 to 25 inches. The females tend to be a little larger than the males. Its feathered body is white in color on its head, neck, and tail, while the rest of the plumage is dark brown. The bald
eagles bill, talons, and iris are yellow. Young bald eagles plumage is completely dark brown until they reach maturity and gradually gain their white feathers from ages three to five. Although scientists do not
know why they have evolved with the white head, it has been suggested that their conspicuous appearance signifies sexual maturity and that it may help serve as a warning to other animals that they have
occupation and dominance of that territory (Stalmaster, 1987).

Natural History:
The bald eagle was formerly widespread all over North America, but now can be found mainly along the Pacific
Northwest with much more sparse populations through the interior to the East Coast (Stalmaster, 1987). You will find bald eagles near seacoasts, reservoirs, lakes, and streams where fish populations are abundant enough to support them. They prefer large and tall trees with open edges (both broadleaf and pine
needle), so as to have easy accessibility and surveillance of prey (INRIN, 2002). Most often they will choose to roost and nest in the tallest tree in the area that is closest to the water. Tall, dead trees or burned up trees will also make a good spot. During cold seasons they may seek more sheltered trees
or floodplains surrounded by river bluffs for less severe temperatures. The bald eagles aggressive behavior near nest sites indicates that the breeding pair establishes a territory with a specific range where they have monopoly over the resources. The average territory ranges from one to two square
kilometers, and their biggest competitors are other bald eagles due to scarcity of resources and the fact that species of their own kind fill the same niche (Stalmaster, 1987).

The bald eagle primarily feeds on fish, especially
ones small in size that they can swallow whole. They will also eat small mammals, small birds, injured waterfowl, and carrion when abundance of fish is not available (INRIN, 2002). Young bald eagles eat the
same as adults. An interesting adaptation the bald
eagles have acquired for the purpose of becoming great
hunters is their vision. Their eyes occupy most of the
space on their head, have both monocular and binocular
vision, and can see three to four times greater
distance than humans making vision their most
important sense (Stalmaster, 1987). Their feet, talons, and beak are also adapted for efficiently tearing and killing slippery prey.

      Bald eagles are monogamous and establish a pair bond for as long as both partners are alive  (INRIN, 2002).
If one of the eagles dies, then the widow will take a new mate, sometimes in as little as three days. Trio bonds have also been known to form, but this is an anomaly, as well as an adult taking an immature mate
(Stalmaster, 1987). Sexual maturity is reached at ages five to six, and courtship is done by rituals, such as vocal displays, chase displays involving acrobatic aerial dives and rolls, and the most spectacular, roller- coaster flights where the two birds dive at great speed, doing somersaults while talons are locked together, then swinging up again before hitting the ground and repeating the process (Stalmaster, 1987). Copulation only happens after the couple has lived together for awhile, and is initiated by the female
when she goes into a solicitation posture and calls over to the male for service. Copulation lasts from 15 seconds to two minutes, but can occur several times a day (Stalmaster, 1987). They breed from February to
March, and lay two to three eggs in March to April with an incubation period of 35 days. Nests are built in the same spots each year, they build the new on top of the old. Nests are built from sticks, grass, leaves, needles, and herbs (INRIN, 2002). Young stay in nests for up to thirteen weeks and begin flying at that time, however, the adults continue to protect and feed the young. Bald eagles are territorial during all the stages of rearing the eaglets. Other duties include sheltering from weather and guarding from
predators, and both parents take on the responsibilities (INRIN, 2002).

Distribution:
Like most diurnal species in the order Falconiformes, Bald eagles are restricted to a single region (Weidensaul,1996). Their indigenous region is North America with occasional stray birds making it to Siberia and Greenland. Their northern most extent ends with the Arctic tundra, because they need trees for a suitable habitat, while their southern most extent is Baja, and across the southern border of the U.S. (Stalmaster, 1987). Why they are limited to this region is probably best explained by their evolution.
Historically, bald eagles were widespread throughout North America and occupied a variety of habitats, including deciduous forests, redwoods, coniferous forests, coastal cliffs and valleys, as well as inland
cliffs and valleys (Detrich, 1983). Their range and population began to shrink as soon as European settlement kicked in, the pre-1800 population is estimated at a quarter million while current estimates are at about eighty thousand (Weidensaul, 1996). They once existed in every state, but now eighteen states are left with no breeding pairs (Salmaster, 1987). An interesting example of their habitat fragmentation is that in California three quarters of their nests are near reservoirs or dams (Detrich, 1983). Although bald eagles have a continuous distribution within a single region, human impact on their niche has chopped it up and made it more of a disjunct pattern. The restrictions to bald eagle distribution are abundant food supply, which is the most important, but also of importance is the need for old- growth forest and freedom from human disturbances (Stalmaster, 1987). One way of adapting to seasonal change in food supply is migration. Birds that nest in the greater part of Canada during breeding season will migrate south into the states during fall, because lakes and rivers freeze during winter in the harsher climates of their range making food supply dwindle (Stalmaster, 1987). Not all eagles migrate, in the milder, year round climates along the coastlines there are birds that stay stationary throughout the year, as well as birds that only nest there or only winter there. This
range runs from Northern California all the way up the coastline to Alaska, and on the East Coast, practically all of Florida, then a disjunct pattern all the way north to Newfoundland. What is interesting about this distribution is that the breeding range of the southern areas, and the wintering range of the northern areas remain exclusive of each other, except at the much smaller regions along the coasts, where you will also find every non- migratory bird, as well as the biggest population concentrations. As far as particular areas of concentrations, Alaska has the most impressive with a thirty- five to forty- five thousand population of wintering eagles (Stalmaster, 1987). The most important single sight is along the Chilkat River in Southeast Alaska where you will find four thousand eagles feasting on spawning salmon within forty miles of each other in November. The second biggest concentration will be found in British Columbia where over twenty- eight thousand
eagles during winter (Stalmaster, 1987). The largest winter population in the United States is found in Washington with close to fifteen hundred birds, primarily in the protected Olympic National Park. Many bald eagles also winter along the Mississippi River in Missouri and Illinois where you will find up to a
thousand. Other worthy mentions will be at McDonald Creek in Glacier Park, Montana, where over five hundred birds winter, and in Florida where there is over one thousand wintering eagles and the largest population of breeding pairs in the forty- eight contiguous states at over three hundred. As far as
concentrations closest to California, you will find over five hundred in the Klamath Basin of Southern Oregon and Northern California, and small populations of wintering and non- migratory birds in Shasta and Trinity national forests (Stalmaster, 1987).         
                                
Evolution:
All birds, which compose the class Aves, arose from reptiles. Whether they came from crocodilelike thecodonts, which are ancestors of dinosaurs, or from small dinosaurs themselves is still debated in the scientific community (Weidensaul, 1996). The best fossil example of an intermediary phase between reptile and bird comes from the Jurassic period with the species Archaeopteryx, which had feathered wings and tail with reptilian teeth and clawed wings. Archaeopterx died out 65 million years ago at the great extinction, but with dinosaurs out of the way, other bird species that had already evolved were now free to expand during the Tertiary period (Weidensaul, 1996). The first modern raptors, whose diurnal members make up the order Falconiformes, appear in the fossil records from the Eocene epoch about 50 million years ago. From this epoch also evolved the ancestral kites, which made up the family Acciptiridae, and the genetic
framework for the 205 species in that family today (Stalmaster, 1987). Included in this family is the genus Haliaeetus, which the bald eagles, along with all eight sea and fish eagles, belong to. The sea eagles of this genus probably came from the scavenging Asian and Australian kites of genus Haliastur, and the
first sea eagle appears 25 million years ago (Stalmaster, 1987). We do not know how far back bald eagles go, but they have found fossil remains in La Brea tar pits that date back over one million years ago. The eight species under the genus Haliaeetus represent a regional distribution that is almost absolutely exclusive of each other. Plate tectonics probably doesn’t explain this, because the eagles are
great dispersers and continental separation had already taken place before they branched out. What their distribution suggests is adaptive radiation and, because of the similarity of ecological niche, maybe a degree of ammensalism helped evolve the eight separate species.


Other interesting issues:
   
As soon as Europeans settled in the New World and expanded West, the bald eagle population began to decline. Coexistence was achieved to somewhat of a degree in certain areas, such as Florida, but in other areas, primarily the ranching parts of the West, bald eagles habit was lost and they were directly killed, because they were seen as a threat to livestock (Weidensaul, 1996). Their diminished numbers were
recognized by the National government, and the Bald Eagle Protection Act was passed in 1940 which made it illegal to kill, harm, or harness any bald eagles, dead or alive, including eggs, feathers, and nests (USF&WS, 2002). Their recovery after this act passed was short lived, because a new threat shortly after began to kill them, that of the chemical pesticide DDT. DDT began by contaminating their food supply, then gradually started accumulating in the fatty tissue of female eagles, which eventually led toreproductive failure through eggshell thinning (USF&WS, 2002). Things began to turn around in 1972, when DDT was banned, and the Endangered Species Act was passed, where the bald eagle was one of its first members. Due to the chemical ban, expanded efforts to locate and understand them, habitat management plans, protective zones, and season closures, the bald eagles numbers in the 48 contiguous states have risen nine times from the 1963 population (Detrich, 1983 and USF&WS, 2002). They were originally classified as endangered in 1972, but in 1995 were reclassified as threatened, making the bald eagle one of the Endangered Species Acts biggest success stories.
Although, this does not mean that the bald eagle is free from danger, or that it has actually recovered.

The bald eagle is still just as much threatened by the same thing that has always threatened them, that is the continually expanding human alteration of the natural environment.

Bibliography:
-Detrich, Philip J. 1983. “Status of the Bald Eagle in California.” Pg.80-83. In Nancy  Venizebs and Celeste Grijalva,eds. RAPTORS. San Francisco Zoological Society.

-Illinois Natural Resources Information Network (INRIC) / Bald Eagle/ www.inhs.uiuc.edu/chf/pub/ifwis/birds/bald-eagle.html/ 10-10-02.       
                                   
-Stalmaster, Mark V. 1987. THE BALD EAGLE. University Books, New York.

-U.S. Fish and Wildlife Service/Bald Eagle/http.endangerd.fws.gov/i/b/msabOh.html/11-1-02

-Weidensaul, Scott. 1996. RAPTORS: THE BIRDS OF PREY.Lyons and Burford Publishers.

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